A young mystic without a past, Kujaku was born under a dark omen possessed of incredible supernatural powers. Raised by priests, he has learned to use these powers for good. But the evil Siegfried von Mittgard seeks to steal his birthright, and rule the world as the Regent of Darkness. He has dispatched bloodthirsty minions to destroy Kujaku before he can awaken to his destiny. Now, Kujaku must unravel the riddle of his past, before the power within consumes him!

Susanoo, the youngest child of the creation gods Izanagi and Izanami, is a mighty warrior, easily able to defeat tigers with his bare hands. After his mother dies, Susanoo is devastated. Determined to travel to Heaven to find her, he carves a boat out of a tree, and along with his rabbit companion, Akahana, he prepares to cross the ocean. As Susanoo does not know how to reach Heaven, he first visits his brother, Tsukuyomi, leader of the Night Kingdom, and his sister, Amaterasu, leader of the Light Kingdom. While his siblings can warn Susanoo of the dangers he faces, they are unable to stop him or provide him with the directions to Heaven. As Susanoo continues his journey to the ends of the Earth, Orochi, an eight-headed dragon, awakens to stand in Susanoo's path.

Suzuko Homura and Chinatsu Morikawa's victory against the nefarious Bookmaker Kou Satomi has allowed the two girls to finally return to a life of normalcy and seize the chance to move on. However, the peace does not last long, as Kiyoi Mizushima requests Suzuko's help to prevent the next cycle of Selector battles; even after Satomi's defeat, efforts to continue his plans seem to persist. Troubled by lingering doubts and the unresolved situation, Suzuko decides to assist Mizushima with her investigation—unknowingly becoming involved in the escalating fight that will determine the fate of Selectors once and for all.